Israeli Prime Minister Benjamin Netanyahu fired back on Monday night after Iran threatened "to 'raze Tel Aviv and Haifa" to the ground, saying that if Tehran attacks Israel, it will be the last anniversary of the Islamic Revolution that they celebrate.
"I do not ignore the Iranian regime's threats, but I'm not impressed by them, either", Netanyahu said in a taped statement.
"If this regime makes the bad mistake of trying to destroy Tel Aviv and Haifa, it will not succeed", he said.
Speaking from a podium in central Tehran, Iranian President Hassan Rouhani addressed the crowds for almost 45 minutes, lashing out at Iran's enemies - America and Israel - and claiming their efforts to "bring down" Iranians through sanctions will not succeed.
The Jerusalem Post reports that Monday's demonstrations featured hundreds of thousands of Iranian citizens - "s$3 oldiers, students, clerics and black-clad women holding small children" alike - and that some burned USA flags.
Javani, the deputy for the IRGC political bureau, was speaking at a rally marking the 40th anniversary of the Islamic Revolution, when the US-backed monarchy was overthrown and replaced in power by Islamic fundamentalists.
Hundreds of thousands of soldiers, militia members in camouflage fatigues, clerics, and ordinary citizens flocked to the streets of other Iranian cities and towns to commemorate February 11, 1979, when followers of Ayatollah Ruhollah Khomeini ousted a USA -backed monarch, Shah Reza Pahlavi.
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Along with the chants "Death to America" and banners bearing slogans calling for "Death to Israel", Monday's marches were also a backdrop to the military's display of Iranian-made missiles, which authorities showcase every year during anniversary celebrations. 40 years of repression.
Rouhani also vowed Iran would defeat US sanctions, reimposed after President Donald Trump withdrew from Tehran's nuclear accord with world powers past year. "The long-suffering Iranian people deserve a much brighter future", he posted in both English and Farsi.
"Now it's up to the Iranian regime to change its behavior, and ultimately up to the Iranian people to determine the direction of their country", national security advisor John Bolton tweeted on the anniversary of the upheaval.
U.S. President Donald Trump pulled out of Iran's 2015 nuclear deal with world powers previous year and re-imposed sanctions on Tehran, dealing a blow to the country's economy. "They should take this into account".
"The Iranian people have and will have some economic difficulties [due to the sanctions] but we will overcome the problems by helping each other", Rouhani said in his address.
Iran also has vast clout in Iraq, where Major-General Qassem Soleimani, head of the overseas arm of the Revolutionary Guards, was frequently photographed guiding Shi'ite militias in the war against Sunni Islamic State militants.